I find this fascinating. LUCA clearly predates some pretty core, basic metabolic capabilities, and yet nucleotide metabolism is complete. This seems to indicate that LUCA and its progenitors were confined to their hydrothermal vents and that the cells could not survive beyond due to the missing metabolic networks.

I wonder if LUCA was not a single cell, but many different populations in different hydrothermal vents, exchanging genetic material wrapped by accident in the remains of the cell membranes as the host cells die and are ejected from their vents.

Perhaps the missing post-LUCA metabolic enzymes evolved by accident from proteins recruited simply to concentrate the required metal ions that are scarcer and ever more diluted the farther from the black smokers you get. Free living was merely a happy bonus effect.