This is exactly why my comment goes over the head of people who cry just get the basic boards. No, this is why the basic boards for $100 don't cut it. You now need to dive into the technical data and realize that the $100 board seems like a deal for a reason, and suddenly the $300+ category is your only option if you want to get a PC that doesn't run on fake specs.

I'm just struggling to figure out how many people actually need the PCIe lanes for anything more than GPUs and storage, though.

Like, what are you actually connecting your desktop to?

The only reason laptops depend on Thunderbolt is because they have limited internal expansion and need high performance external I/O.

If you need more things than gaming boards offer then obviously you have very advanced needs and can go pay for a workstation board, something like an sTR5 socket Threadripper board.